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Megasecoptera: morphology and taxonomy of newly discovered specimens from Upper Carboniferous of northern China (Insecta: Palaeodictyopteroida)
Pecharová, Martina ; Prokop, Jakub (advisor) ; Godunko, Roman (referee)
The extinct insect group of Palaeodictyopteroida (Insecta: Palaeoptera) comprised mainly phytophagous species and occurred from Upper Carboniferous to Upper Permian. Megasecoptera is one of several orders of belonging to this group. Although, the order Megasecoptera comprised over 20 described families, morphology of some body structures are insufficiently studied. The present thesis deals with evaluation of new material comprising the extensive set of 76 fossil insect specimens from the Upper Carboniferous (Bashkirian) of northern China. The fossils having excellent state of preservation of the wings and other body structures provide new insights concerning the external morphology of Megasecoptera. All studied specimens were attributed based on wing venation pattern into two known megasecopteran genera within families Brodiopteridae and Sphecopteridae. The aim of the present thesis is examine the morphology and variability of wing venation of two newly proposed species Brodioptera sp. n. and Cyclocelis sp. n. from site in northern China. In addition the following methods of geometric morphometrics based on landmarks were used for comparison of venational characters: a comparison of centroid size, procrustes analysis, principal component analysis and thin plate spline. Variability of wing venation...
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Morphology and evolution of selected groups of Palaeodictyopterida (Insecta: Palaeoptera)
Pecharová, Martina ; Prokop, Jakub (advisor) ; Hodunko, Roman (referee) ; Sroka, Pavel (referee)
Palaeodictyopterida is remarkable insect superorder, which formed a significant part of the diversity of upper Palaeozoic insects, but disappeared by the end of the Permian. The main synapomorphy of the superorder is the piercing-sucking mouthparts in the form of a rostrum consisting of five styles. This rostrum was probably used to pierce on plant tissue and for the juice sucking. The same type of mouthparts shared by adults was present also in larvae of Palaeodictyopterida. The external copulatory organs of the superorder members was also showed some morphological interests. The male genitalia consist of a pair of gonostyli and two penial lobes, similarly to the genitalia of recent Ephemeroptera. The female genitalia of Palaeodictyopterida are developed in a form of the ovipositor that can be compared with the endophytic ovipositor of some recent Odonata. This morphological features support placement of Palaeodictyopterida as sister group of Odonatoptera + Panephemeroptera. The main aim of the work was to describe new representatives of the order Megasecoptera, the second largest group of Palaeodictyopterida. Wing venation of Megasecoptera exhibits a reduction of the longitudinal and transverse veins in comparison with the order Palaeodictyoptera. Other body structures were examined mainly in the...
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Palaeodictyoptera: morphology of immature wings from the Upper Carboniferous of Poland
Tippeltová, Zuzana
Insect wings are very specific and unique structures in animal kingdom. Wing morphology is a result of long-standing complicated evolutionary process and until recently the way how the wings have evolved is not completely clarified. The flight ability is one of the most important event in insect history because it allows them to exploit new habitats, escape from predators or find the sexual partner. Here we present the newly discovered material consisting of Palaeodictyoptera immature wings from the Upper Carboniferous (Westphalian A) of Poland. This order became extinct in the end of Permian, however during the Late Paleozoic was remarkably diversified. Until recently, number of adult palaeodictyopterans have been described, however the immature stages are relatively unknown due to lack of suitable fossils. Immature wings present in this thesis have undoubtedly palaeodictyopterous affinities with atribution within superfamilies Breyeroidea and Homoiopteroidea. However, their familial assignment into Breyeriidae (morphotype A) and Homiopteridae (morphotype B) based on fore wing venation characters is not definite because of wing venation limits in early ontogenetic stages. The aim of the present work is a complex description of 14 new palaeodictyopteriids immature wings, and to point out certain...
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Morphology and evolution of selected groups of Palaeodictyopterida (Insecta: Palaeoptera)
Pecharová, Martina ; Prokop, Jakub (advisor) ; Hodunko, Roman (referee) ; Sroka, Pavel (referee)
Palaeodictyopterida is remarkable insect superorder, which formed a significant part of the diversity of upper Palaeozoic insects, but disappeared by the end of the Permian. The main synapomorphy of the superorder is the piercing-sucking mouthparts in the form of a rostrum consisting of five styles. This rostrum was probably used to pierce on plant tissue and for the juice sucking. The same type of mouthparts shared by adults was present also in larvae of Palaeodictyopterida. The external copulatory organs of the superorder members was also showed some morphological interests. The male genitalia consist of a pair of gonostyli and two penial lobes, similarly to the genitalia of recent Ephemeroptera. The female genitalia of Palaeodictyopterida are developed in a form of the ovipositor that can be compared with the endophytic ovipositor of some recent Odonata. This morphological features support placement of Palaeodictyopterida as sister group of Odonatoptera + Panephemeroptera. The main aim of the work was to describe new representatives of the order Megasecoptera, the second largest group of Palaeodictyopterida. Wing venation of Megasecoptera exhibits a reduction of the longitudinal and transverse veins in comparison with the order Palaeodictyoptera. Other body structures were examined mainly in the...
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Palaeodictyoptera: morphology of immature wings from the Upper Carboniferous of Poland
Tippeltová, Zuzana
Insect wings are very specific and unique structures in animal kingdom. Wing morphology is a result of long-standing complicated evolutionary process and until recently the way how the wings have evolved is not completely clarified. The flight ability is one of the most important event in insect history because it allows them to exploit new habitats, escape from predators or find the sexual partner. Here we present the newly discovered material consisting of Palaeodictyoptera immature wings from the Upper Carboniferous (Westphalian A) of Poland. This order became extinct in the end of Permian, however during the Late Paleozoic was remarkably diversified. Until recently, number of adult palaeodictyopterans have been described, however the immature stages are relatively unknown due to lack of suitable fossils. Immature wings present in this thesis have undoubtedly palaeodictyopterous affinities with atribution within superfamilies Breyeroidea and Homoiopteroidea. However, their familial assignment into Breyeriidae (morphotype A) and Homiopteridae (morphotype B) based on fore wing venation characters is not definite because of wing venation limits in early ontogenetic stages. The aim of the present work is a complex description of 14 new palaeodictyopteriids immature wings, and to point out certain...
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Palaeodictyoptera: morphology of immature wings from the Upper Carboniferous of Poland
Tippeltová, Zuzana ; Prokop, Jakub (advisor) ; Nel, Andre (referee)
Insect wings are very specific and unique structures in animal kingdom. Wing morphology is a result of long-standing complicated evolutionary process and until recently the way how the wings have evolved is not completely clarified. The flight ability is one of the most important event in insect history because it allows them to exploit new habitats, escape from predators or find the sexual partner. Here we present the newly discovered material consisting of Palaeodictyoptera immature wings from the Upper Carboniferous (Westphalian A) of Poland. This order became extinct in the end of Permian, however during the Late Paleozoic was remarkably diversified. Until recently, number of adult palaeodictyopterans have been described, however the immature stages are relatively unknown due to lack of suitable fossils. Immature wings present in this thesis have undoubtedly palaeodictyopterous affinities with atribution within superfamilies Breyeroidea and Homoiopteroidea. However, their familial assignment into Breyeriidae (morphotype A) and Homiopteridae (morphotype B) based on fore wing venation characters is not definite because of wing venation limits in early ontogenetic stages. The aim of the present work is a complex description of 14 new palaeodictyopteriids immature wings, and to point out certain important...
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Megasecoptera: morphology and taxonomy of newly discovered specimens from Upper Carboniferous of northern China (Insecta: Palaeodictyopteroida)
Pecharová, Martina ; Prokop, Jakub (advisor) ; Godunko, Roman (referee)
The extinct insect group of Palaeodictyopteroida (Insecta: Palaeoptera) comprised mainly phytophagous species and occurred from Upper Carboniferous to Upper Permian. Megasecoptera is one of several orders of belonging to this group. Although, the order Megasecoptera comprised over 20 described families, morphology of some body structures are insufficiently studied. The present thesis deals with evaluation of new material comprising the extensive set of 76 fossil insect specimens from the Upper Carboniferous (Bashkirian) of northern China. The fossils having excellent state of preservation of the wings and other body structures provide new insights concerning the external morphology of Megasecoptera. All studied specimens were attributed based on wing venation pattern into two known megasecopteran genera within families Brodiopteridae and Sphecopteridae. The aim of the present thesis is examine the morphology and variability of wing venation of two newly proposed species Brodioptera sp. n. and Cyclocelis sp. n. from site in northern China. In addition the following methods of geometric morphometrics based on landmarks were used for comparison of venational characters: a comparison of centroid size, procrustes analysis, principal component analysis and thin plate spline. Variability of wing venation...
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